School of Agriculture, Food and Ecosystem Sciences - Theses
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ItemPhysiological aspects of root growth of Eucalyptus pauciflora, subsp. pauciflora and Eucalyptus nitens( 1997)This thesis examined i) morphological and physiological effects of low soil temperatures on root growth of subalpine Eucalyptus pauciflora Sieber ex Sprengel subsp. pauciflora and montane Eucalyptus nitens (Deane & Maiden) Maiden, ii) determined the variability, and in particular the day/night variability, in root elongation, and iii) explored the physiological basis for such variability. A series of experiments were undertaken with seedlings of E. pauciflora and E. nitens grown at soil temperatures of 3, 7, and 13C, and where seedlings were transferred from one temperature to another. E. nitens grew faster than E. pauciflora at 7 and 13C, but E. pauciflora grew faster than E. nitens at 3C. E. pauciflora always produced greater total and white root length than E. nitens. E. nitens roots browned faster in response to lowering soil temperatures than E. pauciflora. The osmotic potential of the roots decreased with soil temperature, but more so in E. pauciflora than E. nitens. Proline was a prominent osmoregulant in roots of E. pauciflora and arginine in E. nitens roots. It is suggested that E. pauciflora is better adapted than E. nitens to root growth at low soil temperatures because it can keep roots white longer and can maintain lower root osmotic potentials. Root growth of E. pauciflora was examined for 31 months (December 1992 - June 1995) in a mature stand at an elevation of 1545 m on Mt Stirling, Victoria, Australia. Greater night than day root elongation was recorded from eight in situ rhizotrons during the summer and early autumn of 1993. Shoot growth was also monitored during part of this study (April 1994 - June 1995). It was found that root growth commenced in the spring at soil temperatures 5 1.5C, under 550 mm of snow, at least one month before the onset of shoot growth and continued at least two months longer that shoot growth during the autumn. A period of root dormancy for at least one month a year occurred in roots of E. pauciflora. The seasonal variability in root numbers of E. pauciflora appeared to be related mainly to soil temperature and to a lesser extent to soil water content. Moreover, there appeared to be some internal periodicity in root growth which was independent of the external environment on Mt Stirling. Greater night than day root elongation was recorded in seedlings of both eucalypts in a glasshouse. Root elongation rates were greatest in E. nitens, and root elongation of both eucalypt seedlings were greater than that of the mature E. pauciflora on Mt Stirling. The zones of day and night elongation were determined in root marking experiments. Histological studies of the zone of elongation showed that cell division occurred mainly during the day and cell elongation mainly at night. Night root elongation rates were increased by increasing day-time air temperatures, light-period, and light intensity; and by decreasing water stress during the night. The turgor pressure of the root tips was greater during the night than the day. It is suggested that the amount of root growth during the night is determined directly by turgor pressure during the night and indirectly by processes during the day (light duration and intensity, and temperature during the light period) which determine the extent of cell division during the day. A greater rate of cell division during the day will be translated into a greater rate of root elongation, especially in the night.
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ItemAn evaluation of fuel-reduction burning in the dry sclerophyll Wombat State Forest( 1994)Each year, between one and three per cent of the total forested area in Victoria is fuel-reduced as a part of the Department of Conservation and Natural Resources' program to implement its overall fire protection policy. The dry sclerophyll Wombat State Forest has been fuel-reduced since the 1960's, - a practise considered important because of its susceptibility to fire and its highly flammable plant community. This thesis presents the results of an evaluation of fuel-reduction burning in the Wombat State Forest, commencing with a review of forest conditions, principles of fuel-reduction burning, fire behaviour and fire effects. This is followed by observations and measurements in the field to document current practices of fuel-reduction burning, to select an appropriate fire-behaviour prediction model, to identify time-intervals between successive burns and to evaluate the effects of fuel-reduction burning on forest trees. Historical data of wildfires were used to evaluate the effectiveness of fuel-reduction burning in limiting the number, area and losses from wildfires. "Multi-criteria analysis" was then applied to select priority areas for fuel-reduction burning. The fuel-reduction burning program which is conducted 'every year during autumn and spring requires detailed planning and preparation, because it can only be carried out under certain prescriptions for weather, fuel and fire behaviour. Depending on the extent and conditions of each area, either ground or aerial ignition is applied. Large areas ignited from the air often result in variations of fire behaviour and intensity that leave many un-burnt patches within the broad areas of burnt forest. In order to predict fire-behaviour, comparisons were made between the Control Burning Meter, the McArthur Fire Danger Meter and the Fire Behaviour Tables for Western Australia (the Red Book). From a study of small trial plots, it was shown that the fire behaviour model incorporated in the Control Burning Meter for messmate-gum or silvertop forest type provided quite realistic predictions of low-intensity prescribed fires compared with predictions based on the McArthur Forest Fire Danger Meter and the Fire Behaviour Tables for Western Australia. Since its inception, fuel-reduction burning in the Wombat State Forest has decreased the quantity of litter and twigs, but it has had no significant effect on the quantity of humus and coarse fuels. This reduction of flammable fuels has reduced the potential for major fires and the study showed that a fuel-reduction burning cycle of less than five years is required to restrict the build up of fine fuels to acceptable levels. With regard to the impact of fuel-reduction burning, it was shown that it causes scorches on most trees, and that there is a significant relationship between scorch-height and tree diameter for the two dominant species, messmate (Eucalyptus obliqua L'Herit) and peppermint (E. radiata Sieb. ex. DC.). However, because of their different bark types, average scorch height was higher for messmate. Due to the considerable variation in the severity of fire seasons, hence in the frequency and severity of fire, no significant differences were detected in the number of wildfires and the extent of areas burnt, before and after the application of fuel-reduction burning in the Wombat State Forest. However, analysis of the historical fire data led to the conclusion that fuel-reduction burning has achieved its objective of limiting the severity of wildfires. Most of the wildfires occurred on areas that either had never been fuel-reduced or that had been fuel-reduced for more than five years. The costs of fuel-reduction burning have been relatively modest in comparison with the potential losses. Therefore, when properly planned and implemented, fuel-reduction burning is a valuable management tool for protecting forests and wider community values. This study has also provided a general idea on the applicability of "multi-criteria analysis" for identifying priority areas for fuel-reduction burning in the Wombat State Forest, with priority areas being selected on the basis of level of fire hazard (fuel, weather and topographical conditions) and values of particular sites (significant values, timber quality, distance from the nearest township and extent of the area). The result was a ranking of areas according to their priorities for burning.
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ItemAnalysis of growth and yield in uneven-aged, mixed species eucalypts at Mt. Cole State Forest( 1994)Continuous forest inventory (CFI) is an effective method for studying forest changes over time; it provides growth and yield information which forest planners and managers can use for planning and managing forests to meet long-term sustained yield. This thesis reports results from a 30 year series of successive CFI measurement at Mt. Cole State Forest in west-central Victoria, commencing in 1963. With an area of 12,352 ha, Mt. Cole State Forest consists of three sub areas; the northern and southern Blocks totalling 11,250 ha, and the Mt. Lonarch Block comprised of 1,102 ha. Permanent sample plots (CFI plots) were first established in the Mt. Cole Blocks in 1963 and these have been measured on seven occasions, including the most recent 1994 assessment. During this period, various changes to the inventory data-base have occurred, which have impacted on the analysis and interpretation of the data. Since 1983 the forest area of the Mt. Cole Blocks have been classified according to land use and management zoning, resulting in a reduction of the net productive area available for sawlog production to 2,758 ha. This area has also been stratified into four homogenous areas (strata) and the number of CFI plots has been reduced from an initial 114 to 66 plots. In the latest (1994) remeasurement, the 66 permanent sample plots for the four strata of the Northern and Southern Mt. Cole Blocks were rechecked and the data has been analysed to determine current growth and yield, and to estimate future volume growth and an available cut for the forest. The above analyses show that the structure and yield of the forest have changed from time to time, with a trend towards decreasing levels of available growing stock over time. The current (1994) growing stock levels of the Mt. Cole Blocks are approximately 80,676 m2�27% (P=0.95) or 28.6�3.7 m2 /ha for basal area, and 237,684 m3�36% (P=0.95) or 83.7�13.9 m3/ha for 'sawlog plus potential sawlog volume'. The current available sawlog volume is approximately 177,672 m3�44% (P=0.95) or 63.5�12.5 m3/ha. Additional analyses of periodic and annual growth rates of the growing stock for each stratum and the whole forest (Mt. Cole Blocks) show that the average diameter increment (underbark) of the forest is approximately 0.51�0.03 cm/yr; the net annual increment including ingrowth (Gn+i) of the growing stock in the Mt. Cole Blocks is approximately 0.63�0.11 m2 /ha/yr for basal area, 2.16�0.75 m3 /ha/yr for sawlog, and 1.78�0.68 m3/ha/yr for sawlog plus potential sawlog. The net annual changes in growing stock levels (Gd) are approximately 0.11�0.29 m2 /ha/yr for basal area, -0.15�1.52 m3 /ha/yr for sawlog, and -0.65�1.52 m3/ha/yr for sawlog plus potential sawlog. Models for predicting the current and future growing stock levels were developed using multiple regression based on the three parameters of initial basal area (BO), initial volume (Yo), and the interval of time between successive measurements (t ). Regulating forests to obtain long-term sustained yields requires a continuous flow of information on growth and yield and the development of comprehensive management strategies based on this information. In this study, such growth and yield information have been used to determine an annual available cut for the Mt. Cole Blocks of approximately 5,364�1,854 m3/yr.
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ItemThe occurrence of brittleheart in Eucalyptus regnans and its effect on various wood properties( 1990)This project is mainly concerned with a description of anatomical and ultrastructural characteristics of cell wall deformations in brittleheart of E. regnans, development of methodology for quantification of percentage of broken fibre pieces (PBFP) in E. regnans, and physical and anatomical properties of E. regnans in relation to the occurrence of brittleheart. Two 1939 regrowth E. regnans butt logs and one mature growth E. regnans butt log removed from a tree approximately 120 years old were used in the study. The anatomical and ultrastructural characteristics of cell wall deformations were examined with bright field, polarized light, and scanning electron microscopy. The microscopic compression lines were found to consist of slip planes, minute compression failures, corrugations, and buckles. The width of microscopic compression lines along the longitudinal fibre axis ranged from one single fold in the cell wall up to 7 times the average fibre width. The length of the microscopic compression lines ranged from ones which only crossed a few fibres to ones which crossed up to 200 fibres. The severity of microscopic compression lines appeared to be dependent on the type of cell wall deformations and degree of compaction in the deformed zone. In the methodology studies it was found that for a pulp sample prepared from a 2 mm by 2 mm by 15 mm wood chip approximately 400 cellular elements in 8 out of 16 areas on a slide needed to be counted in order to obtain PBFP with less than 10% error. For macerated thin sections, all the cellular elements on 2 slides each carrying about 600 fibres and fibre pieces needed to be counted. A highly significant correlation was found between the length of microscopic compression lines per unit area and the microscopically determined PBFP. A highly significant correlation was found between the theoretical PBFP and the microscopically determined PBFP. These findings confirm that maceration of wood containing cell wall deformations results in broken fibres and verifies the validity of the maceration technique for quantifying the amount of microscopic compression lines. PBFP was found to increase with macerating time. A maceration time around 10 hours appears sufficient to cause fibres having cell wall deformations to break but longer times cause fibres without cell wall deformations to also break due to overmaceration. A significant relationship was found between PBFP determined after 5 hours and after 10 hours of maceration at the same temperature. It was found that parameters extracted or derived from cell length distributions produced by the Kajaani FS-200 may be used to determine the amount of fibre fragments in pulp samples. These parameters were the high peak, the length weighted average, and the mass weighted average of the cell length distributions for pulps which had PBFP greater than 10. Based on the maceration technique, brittleheart was detected in both the mature and the 1939 regrowth logs. Brittleheart was more severe and occupied a larger area in the mature growth than in the regrowth wood. The PBFP was found to be mostly below 5 in the regrowth wood although relatively high PBFP values of 21 and 30 were observed. PBFP values as high as 85 was found in the mature growth wood. In general, PBFP was found to be higher nearer the pith, decreasing toward the bark, and dropping to zero before reaching the two-thirds theoretical point where the stress is assumed to be 0. A large circumferential variation in PBFP was observed in four adjacent growth rings of the mature log. A large variation in PBFP was also found within a volume of 1000 mm^3 for both the mature and the 1939 regrowth wood. The earlywood PBFP was found to be significantly higher than the latewood PBFP for the mature and regrowth logs. A total of 72 green and 132 12% MC Izod specimens were prepared from the two 1939 regrowth logs and tested for impact strength. The mean Izod value was found to be 9.9 ft.lb for the 132 12% MC Izod specimens and 9.2 ft.lb for the 72 green Izod specimens. For side-matched Izod specimens, the mean Izod value was found to be 9.2 ft.lb at green and 9.7 ft.lb at 12% MC and the mean for the 12% MC specimens did not reflect the expected increase in strength with moisture loss. Excessively low Izod values (eg. 1.8 ft.Ib) were found in the 12% MC Izod specimens located near the pith. For 76 12% MC Izod specimens, their PBFP, fibre length, and specific gravity were also measured. Significant relationships (p=0.01) were found between Izod values and specific gravity, PBFP, and fibre length for these 76 12% MC Izod specimens. Specific gravity and fibre length positively contribute to the impact strength whereas PBFP negatively affects the impact strength. Brash-break specimens showed a low mean Izod value, a low mean specific gravity, the presence of brittleheart, and a short mean fibre length.